Král, D. ; Bezděk, A. (2016). “Tribe Scarabaeini Latreille, 1802”. In Löbl, I. & Löbl, D.. Catalogue of Palaearctic Coleoptera, Volume 3. Revised and Updated Edition. Scarabaeoidea – Scirtoidea – Dascilloidea – Buprestoidea - Byrrhoidea. Leiden, The Netherlands: Brill. pp. 204–207. doi : 10.1163/978...